Small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark cream, gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; compound; pinnate; with a terminal leaflet; leaflets opposite, or leaflets alternate; all of the same type; not modified into spines; coriaceous; gland-dotted; very asymmetric at the base; not dissected; wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present. Stalk of blade absent to less than a quarter the length of the blade; winged (narrowly), or not winged; attached to base of leaf blade; not swollen. Stipules absent (be careful not to confuse with lower leaflets which are stipule like). Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; axillary; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across; somewhat asymmetric; with male or female parts raised on a column; with the female parts of the flower raised on a stalk. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked (tapering to base); entire. Stamens 10; filaments present; free of each other; free of the perianth; of different sizes; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 5(-6). Locules 5(-6). Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaflets of different sizes, the basal ones like stipules; petioles narrowly winged; fruits mucilage filled; seeds numerous and densely scaly. Taxon images: merrill.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.