Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like, or box like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; laterally placed; encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences represented by solitary flowers only; terminal. Flowers bisexual; stalked; to about 3 cm long or across to more than 10 cm long or across; round; with male or female parts raised on a column, or not raised on a column; with the female parts of the flower raised on a stalk. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 9-21. Stamens 100 (many); filaments present; free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; not awned; dehiscing by long slits. Ovary superior. Carpels separate, or partially joined, by base; (2-)100. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit follicular; fleshy, or non-fleshy; multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Stipules large and amplexicaul; seeds hang from fruiting bodies on the funicles; 4 ovules per carpel. Taxon images: mangliet.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.