Small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate present; white, or yellow; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; of different sorts on a single twig, or all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; leaf-opposed, or on the trunk or branches; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 2; partially joined; persistent in fruit, or not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles solitary. Stigmas 2 lobed; distinctive; disk-like. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-4 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Small trees; leaves with recurved margins; hypsophylls present, sometimes persistent; sepals at first fused, but then splitting into two valves. Taxon images: mammea.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.