Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark white, cream, gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; disk-shaped, or star-like. Stinging hairs absent. Mature twig indumentum present; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3; free; not fleshy; deciduous; not stalked; entire. Stamens 3; filaments present; free of each other; free of the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined, or partially joined, by base; 3. Locules 3. Styles solitary. Stigmas 3 lobed; not distinctive. Fruit follicular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 3 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Part covered in scales or stellate hairs; flowers in small globose clusters; flowers unisexual and in parts of 3.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.