Small trees to shrubs. Buttresses absent. Spines present, or absent; on trunk, or on branches. Stilt roots absent. Bark green, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; star-like. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple, or compound; with three leaflets, or pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; not dissected; wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; winged, or not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; encircling the twig; scale-like; not fringed; large, or small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; leaf-opposed; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present, or absent; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; all joined (base); not fleshy; deciduous; not stalked; entire. Stamens 4-5; filaments present; joined; joined to the perianth; all equal in size; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-4. Locules 4-8. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4-8 per fruit; less than 5 mm; wingless; with folds like marble, in cross- section; aril absent; endospermic.
Treelets or shrubs; sometimes with thorns; leaves compound with generally serrate margins; leaflet tertiary venation ladder-like; stipules on petiole. Taxon images: leea.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.