Emergent trees to large trees. Buttresses present; tall. Spines absent. Stilt roots absent. Bark white, cream, gray, black; not peeling in strips. Slash orange, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral, or two-ranked; not scale-like; compound; pinnate; with a terminal leaflet; leaflets alternate; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible, or not visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; free of each other; free of the perianth; all equal in size; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by pores, or dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit a samara; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged. Seeds 1 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Canopy trees; bark either dark or silvery white; buttresses tall and often spreading; flowers minute; seed flat and prominently winged. Taxon images: koompass.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.