Large trees to shrubs. Buttresses present; short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; yellow; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open, or closed; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; terminal, or axillary; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4; partially joined; persistent in fruit, or not persistent in fruit (rare); growing slightly after flowering. Epicalyx absent. Petals present; 4; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; (3-)4. Locules 1. Styles absent. Stigmas 2-4 lobed; distinctive; head shaped. Fruit capsular; non-fleshy; not multiple; splitting; enclosed in, or sitting on, the fleshy perianth; wingless. Seeds 1-4 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaves with distinct intercostal veins; gland dots present, surrounded by reticulations; anthers often with glands; fruit oten capsular, thinly woody and 1-4 seeded, surrounded by calyx. Taxon images: kayea.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.