Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark red, gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible, or not visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; joined across twigs; encircling the twig; scale-like, or leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5(-6); partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5(-6); free; not fleshy; deciduous; not stalked; deeply cut in two, or fringed like a comb. Stamens 100 (many); filaments present; free of each other; joined to the perianth; of different sizes; polystemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 3. Locules 1. Styles solitary. Stigmas 3 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Mangrove tree without pneumatophores; trunk with swollen conical base; leaf apex blunt to rounded; petals bilobed with a long bristle between the lobes, and each lobe fringed. Taxon images: kandelia.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.