Small trees, or shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; fully joined; not joined together and forming a lid; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; free of each other; free of the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels partially joined, by styles; 2. Locules 2. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit; barely visible; wingless; uniform in cross-section; aril absent; endospermic.
Spiral toothed leaves with ladder-like tertiary venation; inflorescences long racemes; prominent disk surrounding two carpels joined by the style. Taxon images: itea.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.