Large trees to shrubs. Buttresses present, or absent; tall. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash yellow, orange, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present, or absent; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; not stalked, or stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present, or absent; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-12; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-12; free; not fleshy; deciduous; not stalked; entire. Stamens 2-12; filaments present; free of each other; free of the perianth, or joined to the perianth; all equal in size; anisomerous, or isomerous, or diplostemonous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary superior, or partly inferior. Carpels joined; 2-7. Locules 1. Styles solitary, or free; 2-7. Stigmas 1-7 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; enclosed in, or sitting on, the fleshy perianth, or not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-5 per fruit; barely visible, or less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Petiole not kneed; flowers in spikes or racemes; fruits seated within the persistent calyx. Taxon images: genhomal.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.