Small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark cream; not peeling in strips. Exudate present; white; flowing, or spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy. Venation palmate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade; not winged; attached to surface of leaf blade, attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands present, or absent. Hairs on mature leaf absent.
Inflorescences present; terminal; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant, or with male and female flowers on different plants; not stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 2; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 6-10; filaments present; free of each other; free of the perianth; all equal in size; triplostemonous, or polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles free; 2. Stigmas 1 lobed; not distinctive. Fruit capsular; fleshy; not multiple; splitting, or not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Small trees; white latex from cut parts; leaves as long as broad with a delicate appearance; flowers in terminal spikes; fruits bilobed.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.