Small trees to shrubs. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; star-like. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; compound; pinnate; without a terminal leaflet; leaflets opposite, or leaflets alternate; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; winged, or not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; terminal, or axillary, or on the trunk or branches; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; stalked; entire. Stamens 5-7; filaments present; free of each other; free of the perianth; all equal in size; isomerous, or number of stamens not a multiple of the adjacent perianth whorl. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2(-4). Locules 2(-4). Styles solitary. Stigmas 2 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm; wingless; uniform in cross-section; aril present; non-endospermic.
Stellate hairs present; sepals and petals long; fruits 2-3 lobed; seeds 1 per lobe, covered by an aril. Taxon images: genharpu.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.