Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark white, gray, brown; not peeling in strips. Slash white, yellow. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not visible. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4; partially joined; persistent in fruit; growing slightly after flowering (forming a large skirting below the fruit). Epicalyx absent. Petals present; 4, or 6-8 (4 male, 6-8); all joined; not fleshy; deciduous; not stalked; entire. Stamens 4; filaments present; joined; joined to the perianth (sitting on the corolla); all equal in size; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3. Locules 1. Styles solitary. Stigmas 3 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; enclosed in, or sitting on, the fleshy perianth; winged. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaves distichous; intramarginal vein present; tertiary veins invisible; monoecious; calyx accrescent, forming a large skirt below the fruit.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.