Guettarda

Family Rubiaceae. Order Rubiales.

Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.

Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; joined across twigs; encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; sparse.

Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across to to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present. Perianth present; with distinct calyx and corolla whorls. Sepals 0, or 5 (lobes barely visible); partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; (4-)6-8(-9); all joined; not fleshy; deciduous; not stalked; entire. Stamens 4-9; filaments present; free of each other; joined to the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2. Locules 4-9. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4-6 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.

Seashore tree; inflorescences on long peduncles; stigma slightly exserted; ovary 4-9 celled. Taxon images: guettard.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.