Small trees to shrubs, or climbers. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; disk-shaped, or star-like. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis waxy, or not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; fringed, or not fringed; small; not persistent. Domatia present, or absent; pit-like; scattered along midrib. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts raised on a column, or not raised on a column; with the male parts of the flower raised on a stalk. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present, or absent; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary superior. Carpels joined. Locules 1-5. Styles solitary. Stigmas 3 lobed; not distinctive. Fruit capsular, or drupaceous; fleshy; not multiple; splitting, or not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-8 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves trinerved; tertiary veins ladder-like; petals shorter than sepals; fruits generally bright orange and fuzzy, sharp tasting. Taxon images: gengrewi.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.