Gmelina

Family Verbenaceae. Order Lamiales.

Small trees to shrubs. Buttresses absent. Spines present; on branches. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs present, or absent; disk-shaped. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.

Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; dissected, or not dissected; pinnately lobed; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.

Inflorescences present, or represented by solitary flowers only; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across to to about 10 cm long or across; with one plane of symmetry; with male or female parts not raised on a column. Hypanthium absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; partially joined; not persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; 4-5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 4; filaments present; free of each other; joined to the perianth; of different sizes; anisomerous, or isomerous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 4. Locules 4. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.

Axillary spines present; flowers large, yellow or orange; fruit a drupe with one or two big seeds; leaves with glands. Taxon images: gmelina.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.