Large trees to shrubs. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots present, or absent. Bark red, gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; colorless; spotty; blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein absent. Stalk of blade absent to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts raised on a column, or not raised on a column; with the female parts of the flower raised on a stalk. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals (4-)5(-8); fully joined; joined together and forming a lid; not persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals present; (4-)5(-8); free; not fleshy; persistent (enlarged), or deciduous; not stalked; entire. Stamens (4-)5(-7), or 10, or 100 (many); filaments present; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous, or polystemonous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged (petals), or wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Bark often with blackened areas caused by sap; sap extremely irritating; pedicel swollen; calyx calyptrate; petals sometimes accrescent; floral axis sometimes elongated; disk absent; in fruit the style terminal or lateral. Taxon images: gengluta.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.