Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; ladder like to box like. Intramarginal vein absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; joined across twigs; encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences represented by solitary flowers only; axillary. Flowers bisexual; about 0.5 cm long or across to to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; 5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; free of each other; joined to the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Veins prominent, the smaller ones box-like; flowers single, sometimes in pairs; fruit with enlarged persistent calyx; seeds large. Taxon images: gengarde.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.