Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite, or two-ranked; scale-like, or not scale-like; simple; all of the same type; not modified into spines; leathery; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation trinerved. Secondary veins open; not prominent, but visible. Smallest veins not visible. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; not stalked; with the parts barely visible to the eye; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with all whorls looking like tepals. Tepals 4-5. Stamens 4-5; filaments present; free of each other; free of the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles absent. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; enclosed in, or sitting on, the fleshy receptacle; wingless. Seeds 1 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Taxon images: exocarp.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.