Small trees to shrubs. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; with small to large teeth on margin (gland on each tooth). Epidermis not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like to ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at both ends. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls, or intergrading from sepals to petals. Sepals 5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present, or absent; 5; free; not fleshy; deciduous; not stalked; fringed like a comb (apically). Stamens 5; filaments present; free of each other; free of the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3(-4). Locules 1. Styles solitary. Stigmas 1-3 lobed (1 or 3); not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-6 per fruit; less than 5 mm; wingless; uniform in cross-section; aril present (scarlet); endospermic.
Leaves with toothed margins; seeds with a scarlet aril; fruit splitting. Taxon images: eryflac.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.