Large trees to small trees. Buttresses present; tall. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash cream, orange, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present, or absent; white; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs present; star-like. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation palmate. Secondary veins open; prominent. Smallest veins prominent; ladder like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade to more than half the length of the blade; not winged; attached to surface of leaf blade, attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands present, or absent. Hairs on mature leaf present; dense.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; with the parts barely visible to the eye; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3-4; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 6-10; filaments present; joined (base); free of the perianth; all equal in size; diplostemonous. Anthers 4 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-6. Locules 2-6. Styles absent. Stigmas 2-6 lobed; distinctive; disk-like. Fruit capsular; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Trees sometimes buttressed; inner bark cream colored with orange flecks; leaf base heart shaped, sometimes peltate; venation palmate. Taxon images: endosp.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.