Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray to brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to surface of leaf blade, attached to base of leaf blade; swollen; swollen at one end, or swollen at both ends. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense.
Inflorescences present; axillary; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; free; not fleshy; deciduous; not stalked; entire. Stamens 8, or 10; filaments present; joined; free of the perianth; of different sizes; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 2 lobed; not distinctive. Fruit follicular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril present; endospermic.
Leaves distichous and jointed at base; stamens of 2 different sizes; fruits densely hairy; seed with a yellow-orange arillode at base. Taxon images: genelli.gif ellipant.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.