Large trees to shrubs. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash gray, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; star-like. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves opposite, or spiral; not scale-like; compound; pinnate; without a terminal leaflet, or with a terminal leaflet; leaflets opposite, or leaflets alternate; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; winged, or not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present, or absent; scattered across lamina. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary, or on the trunk or branches; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3-5(-6); free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3-5(-6); free, or all joined; not fleshy; deciduous; not stalked; entire. Stamens 6-10(-13); filaments present; joined; free of the perianth, or joined to the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-5. Locules 2-5. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2-10 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril present, or aril absent; non-endospermic.
Leaves opposite of spiral; leaflets generally glabrous. Taxon images: gendysox.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.