Emergent trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash yellow, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal; on a branched axis; specialized inflorescence units present; specialized inflorescence units corymbs; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-8; free, or partially joined; persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; 4-8; free; not fleshy; deciduous; stalked (short); lobed, or entire. Stamens 12-100 (many); filaments present; free of each other; joined to the perianth (calyx); all equal in size; diplostemonous, or triplostemonous, or polystemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary partly inferior. Carpels joined; 4-8. Locules 4-8. Styles solitary. Stigmas 4-8 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Large trees with or without buttresses; leaves simple and opposite; stipules absent; inflorescences terminal corymbs; 4-8 calyx lobes; fruit a capsule with numerous small seeds; seeds "tailed" be extended testa. Taxon images: duabanga.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.