Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash cream. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; very asymmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; fringed, or not fringed (fringed in Drypetes eriocarpa); small; not persistent, or persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary, or on the trunk or branches; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-6; free; not persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals absent. Stamens 4-100 (many); filaments present; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous, or triplostemonous, or polystemonous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-3. Locules 1-3. Styles absent. Stigmas 2 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-3 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaves distichous with strongly asymmetric bases; fruit a leathery drupe; style absent and stigma bilobed, appearing like a bow-tie. Taxon images: drypetes.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.