Large trees to shrubs. Buttresses present, or absent; tall. Spines absent. Stilt roots present, or absent. Bark red, gray, brown; not peeling in strips. Slash red, gray, brown. Vertical stripes in the inner bark or wood present. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; winged, or not winged; attached to base of leaf blade; not swollen. Stipules present, or absent; free from one another, or joined; laterally placed; encircling the twig, or not encircling the twig; leafy; not fringed; large, or small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; terminal, or axillary (1 sp.); on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 10 cm long or across to more than 10 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals (4-)5(-6); free; persistent in fruit, or not persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present, or filaments absent; joined, or free of each other; free of the perianth; all equal in size, or of different sizes; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by pores (usual), or dehiscing by long slits. Ovary superior. Carpels partially joined, by base; 4-20. Locules 1. Styles free. Stigmas 1 lobed; not distinctive. Fruit capsular; fleshy; not multiple; splitting; enclosed in, or sitting on, the fleshy hypanthium, or not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril present, or aril absent; endospermic.
Inner bark brown red; trunk of tree often "hisses" when cut; prominent leaf scars; flowers large; petals yellow or white; stamens numerous; fruit often star shaped and fleshy. Taxon images: gendille.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.