Large trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark white, cream, gray, brown; not peeling in strips. Slash yellow, orange, brown. Vertical stripes in the inner bark or wood present, or absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense.
Leaves whorled; 3-5 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery, or paper-like; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules absent. Domatia present; pit-like; scattered along midrib. Basal glands present, or absent. Hairs on mature leaf present; dense.
Inflorescences present; terminal; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across, or to about 10 cm long or across; with one plane of symmetry; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 4; filaments present; free of each other; joined to the perianth; all equal in size; anisomerous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles solitary. Stigmas 2 lobed; distinctive; club shaped. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm, or about 1 cm long; winged; uniform in cross-section; aril absent; non-endospermic.
Twigs lenticillate; leaves simple, entire and in whorls of three; exstipulate; usually with two or more glands at the base of the adaxial leaf surface; 4 stamens, exserted and bending back over the flower; stigma 2- lobed; seeds winged. Taxon images: deplanch.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.