Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; symmetric at the base; dissected, or not dissected; palmately lobed; smooth margined. Epidermis waxy, or not waxy. Venation trinerved. Secondary veins open; prominent to not prominent, but visible. Smallest veins prominent to not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; terminal; on a branched axis; specialized inflorescence units present; specialized inflorescence units umbels; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 0, or 5; partially joined, or fully joined; not joined together and forming a lid; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 5-8; filaments present; free of each other; free of the perianth; all equal in size; isomerous, or number of stamens not a multiple of the adjacent perianth whorl; alternating with the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined. Locules 4-8. Styles solitary, or free; 5. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4-8 per fruit; barely visible to less than 5 mm; wingless; with folds like marble, in cross- section, or uniform in cross-section; aril absent; endospermic.
Leaves usually simple and trinerved at base; calyx sometimes rim-like; no articulation between petiole and blade; ovary 4 or more celled. Taxon images: dendpax.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.