Dacryodes

Family Burseraceae. Order Sapindales.

Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots present, or absent. Bark green, gray, brown; not peeling in strips. Slash cream, yellow, orange, red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present, or absent; white, or colorless; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.

Complex hairs present, or absent; star-like (Dacryodes nervosa). Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.

Leaves spiral; not scale-like; compound; pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; very asymmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at both ends. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.

Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3; free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3; free; fleshy, or not fleshy; deciduous; not stalked; entire. Stamens 6; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; (2-)3. Locules (2-)3. Styles absent. Stigmas 3 lobed; distinctive; disk-like. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.

Leaflet bases strongly asymmetric; petiolules often long and strongly kneed at both ends; stigma almost or actually apical in fruit; fruit coarsely wrinkled when dry, glabrous, not seated on an accrescent calyx. Taxon images: gendacry.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.