Cyathocalyx

Family Annonaceae. Order Magnoliales.

Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark brown, black; peeling in strips. Slash yellow, red. Vertical stripes in the inner bark or wood absent. Odor aromatic. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs present, or absent; star-like. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.

Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.

Inflorescences present, or represented by solitary flowers only; axillary, or leaf-opposed, or on the trunk or branches (Cyathocalyx biovulatus); the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across, or to about 10 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 3; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 6; free; fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments absent; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; not awned; dehiscing by short slits. Ovary superior. Carpels solitary, or separate; 1-100 (many). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm to much more than 1 cm long; wingless; with folds like marble, in cross- section; aril absent; endospermic.

Sepals forming a cupule; petals spoon-shaped; fruits globose to ovoid, sessile. Taxon images: gencyath.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.