Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present; dense, or sparse.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present (usual), or absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present (rudimentary), or absent; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on different plants; stalked (short); about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5(-6); free, or partially joined; persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals absent. Stamens 4-5(-6); filaments present; free of each other; free of the perianth; of different sizes; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base (adnate); not awned; dehiscing by long slits. Ovary superior to partly inferior. Carpels joined; 2-4. Locules 2-4. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible to less than 5 mm; winged; uniform in cross-section; aril absent; non-endospermic.
Leaves opposite, veins prominent and intramarginal vein present; stipules small; fruit a capsule. Taxon images: cryptero.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.