Large trees to shrubs. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark yellow, red, gray, brown; peeling in strips, or not peeling in strips. Slash green, orange, red, brown. Vertical stripes in the inner bark or wood present, or absent. Odor absent, resinous. Exudate present; red; flowing, or spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined, or free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3. Locules 3. Styles free; 3. Stigmas 1 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible, or less than 5 mm; winged; uniform in cross-section; aril absent; endospermic.
Bark reddish in large trees; orange or red exudate from sap; leaves opposite and gland dotted, often with one or more intramarginal veins; seeds numerous in a dehiscent capsule and winged. Taxon images: gencrato.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.