Copaifera

Family Fabaceae. Order Fabales.

Large trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.

Leaves spiral; not scale-like; compound; pinnate; without a terminal leaflet; leaflets opposite, or leaflets alternate; all of the same type; not modified into spines; coriaceous; gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.

Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts raised on a column; with the female parts of the flower raised on a stalk. Hypanthium unknown. Disk present. Perianth present; with distinct calyx and corolla whorls. Sepals 4; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals unknown; unknown; unknown; unknown; unknown. Stamens 10; filaments present; free of each other; free of the perianth; of different sizes; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by short slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit legume-like; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; about 1 cm long; wingless; uniform in cross-section; aril present (2-lobed and enclosing the seed); non-endospermic.

Large tree; leaves glossy, venation delicate; seeds large with bilobed arils. Taxon images: copaifer.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.