Combretum

Family Combretaceae. Order Myrtales.

Small trees to shrubs, or climbers. Buttresses absent. Spines present, or absent; on branches (petiolar). Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs present, or absent; disk-shaped. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.

Leaves opposite, or whorled, or spiral; 3-5 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined (rare). Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent to not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.

Inflorescences present; terminal, or axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round, or somewhat asymmetric; with male or female parts not raised on a column. Hypanthium present. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; free; not fleshy; deciduous; not stalked; entire. Stamens 8, or 10; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2-5 (reported for family). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit a samara; fleshy (rare), or non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.

Usually climbers in low frequently flooded forest; fruit with 4-5 wings. Taxon images: combretu.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.