Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots present, or absent. Bark gray, brown; not peeling in strips. Slash yellow, red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; disk-shaped. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3-4; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3-4; free; not fleshy; persistent, or deciduous; not stalked; fringed like a comb, or entire. Stamens 6, or 8; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 3(-4). Locules 3-4. Styles free; 3(-4). Stigmas 1 lobed; not distinctive. Fruit a samara; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Stipules absent; leaves alternate; ovary triangular; petals entire to lacinate; styles 3(-4) and free; fruits three winged. Taxon images: gencomb.gif combret.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.