Emergent trees to large trees. Buttresses present; tall. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; disk-shaped. Stinging hairs absent. Mature twig indumentum present; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like to leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense.
Inflorescences present; axillary, or on the trunk or branches; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx present. Petals present; 5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 20(-100) (many); filaments present; joined; joined to the perianth; all equal in size; polystemonous. Anthers 1 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 5. Locules 5. Styles solitary. Stigmas 5 lobed; distinctive; disk-like. Fruit capsular; fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm, or about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril present; non-endospermic.
Prominent buttresses; parts densely covered by silvery scales; petioles slender, kneed; epicalyx 3 lobed; petals shorter than sepals; anthers unilocular; fruit splitting about 2/3 length before falling; seeds black with a thin cream colored aril. Taxon images: gencoelo.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.