Small trees to shrubs, or climbers. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; disk-shaped. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves opposite, or whorled; 3-5 per whorl; not scale-like; simple; of different sorts on a single twig, or all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; very asymmetric at the base, or symmetric at the base; dissected, or not dissected; pinnately lobed; smooth margined to with small to large teeth on margin. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units present, or absent; specialized inflorescence units corymbs; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across to to about 10 cm long or across; with one plane of symmetry; with male or female parts not raised on a column. Hypanthium absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals present; 5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 4; filaments present; free of each other; joined to the perianth; all equal in size; anisomerous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 4. Locules 4. Styles solitary. Stigmas 2 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds (1-)4 per fruit; barely visible to less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Calyx lobes 5, continuing to grow after flowering. Taxon images: cleroden.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.