Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark green, brown; not peeling in strips. Slash orange, red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; with small to large teeth on margin (glands on teeth). Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; net like to ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across to to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4; all joined (base); not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined (base); free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2(-3). Locules 2(-3). Styles solitary. Stigmas 1 lobed; not distinctive. Fruit capsular; fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; much more than 1 cm long to more than 10 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Trees with clustered spirally arranged leaves, margins toothed, the teeth often with a gland; inflorescences terminal panicles; calyx tube hairy. Taxon images: chydenan.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.