Large trees to small trees. Buttresses absent. Spines present; on branches. Stilt roots absent. Bark gray; not peeling in strips. Slash orange, red, brown. Vertical stripes in the inner bark or wood present, or absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at both ends. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present (glandular); the stamens inside the disk, or the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 8-10; filaments present; free of each other; free of the perianth; of different sizes; diplostemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3. Locules 3. Styles free; 3. Stigmas 2 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 3 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Long petioled leaves, margin wavy to serrate; no stellate hairs; sometimes seeds with red, ochre and black layers. Taxon images: cheilosa.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.