Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present (glandular, sometimes adnate to sepals); the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 8; filaments present; joined (base); free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3. Locules 3. Styles free; 3. Stigmas 2 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 6 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Disk glands at the base of the sepals; styles divided in 2; stamens united into a tube below.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.