Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; axillary; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 8-10; filaments present; joined; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary superior. Carpels joined; 2-3. Locules 1. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; fleshy, or non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 10-100 per fruit (many); less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Petiole not kneed; margin usually toothed; flowers small and axillary; fruit ribbed when dry and with the persistent calyx. Taxon images: gencasea.gif gencasgr.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.