Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate, or palmate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; leaf-opposed; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 2; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 6; free; fleshy; deciduous; stalked; entire. Stamens 100 (many); filaments absent; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; not awned; dehiscing by short slits. Ovary superior. Carpels separate; 100 (many). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 3-100 per fruit (many); less than 5 mm; wingless; with folds like marble, in cross- section; aril absent; endospermic.
Crown small and branches at the top of the bole; leaves often chartaceous; all hairs simple. Taxon images: canan.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.