Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery, or paper-like; gland-dotted; symmetric at the base; not dissected; smooth margined to with small to large teeth on margin (usual). Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent to not prominent, but visible. Smallest veins prominent to not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; axillary; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; not stalked, or stalked; to about 3 cm long or across, or to about 10 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5-7(-100) (many); free, or partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5-7(-100) (many); free, or all joined (base); not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined, or free of each other; joined to the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3-5. Locules 3-5. Styles solitary, or free; 3-5. Stigmas 3-5 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Styles normally 3-5, free.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.