Calophyllum

Family Clusiaceae. Order Theales.

Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots present, or absent. Bark yellow, green, orange, red, gray, brown, black; not peeling in strips. Slash red. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; white, or yellow; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.

Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.

Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins closed; not prominent, but visible. Smallest veins not visible. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent.

Inflorescences present; terminal, or axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units present, or absent; specialized inflorescence units umbels; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present, or absent; 2-4; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined, or free of each other (at base); free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; distinctive; disk-like. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.

Bark dark with red or yellow tinges, often with boat shaped fissures; slash with sticky white or dirty yellow exudate; leaves with numerous parallel veins, very close together, perpendicular to the midrib; fruit a one seeded drupe. Taxon images: caloph.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.