Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash cream. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves opposite, or whorled, or spiral; 3-10 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible, or not visible; ladder like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at both ends. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present (glandular). Perianth present; with distinct calyx and corolla whorls. Sepals 3-5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 5; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous, or triplostemonous, or polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3. Locules 2-3. Styles solitary. Stigmas 3 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm, or about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaves with long petioles, kneed at both ends; stamens with glands between their bases. Taxon images: blumeo.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.