Small trees. Buttresses present, or absent; tall. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary, or on the trunk or branches; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; not stalked, or stalked; to about 3 cm long or across, or to about 10 cm long or across, or more than 10 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-6; partially joined; persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals present; 4-6; all joined (base); not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined (base); free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2-4 (2 or 4). Locules 2-4 (2 or 4). Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; about 1 cm long to more than 10 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Fruits generally four angled or winged, especially when young - watch out for great variation in vegetative parts. Taxon images: barring.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.