Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark red, gray, brown; not peeling in strips. Slash cream, gray, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery; not gland-dotted; symmetric at the base; not dissected; with small to large teeth on margin (usually gland tipped). Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; laterally placed; encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant, or with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth absent. Stamens 1-3(-5); filaments absent; all equal in size. Anthers 2 loculed; not awned; dehiscing by long slits, or dehiscing by valves. Carpels solitary; 1. Locules 1. Styles absent. Stigmas 2 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; barely visible, or less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves opposite and glandular-serrate; flowers small and without perianth; aromatic smell to crushed parts; glabrous. Taxon images: genascar.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.