Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; peeling in strips. Slash cream, gray. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; white; flowing, or spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple, or compound; pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; dissected, or not dissected; pinnately lobed, or palmately lobed; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or box like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another, or joined; laterally placed; encircling the twig, or not encircling the twig; leafy; not fringed; large; not persistent, or persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary, or on the trunk or branches; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 2-4. Stamens 1; filaments present; free of each other; free of the perianth; all equal in size; anisomerous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior, or partly inferior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 2 lobed; not distinctive. Fruit drupaceous; fleshy; multiple; not splitting; enclosed in, or sitting on, the fleshy perianth; wingless. Seeds 100 per fruit (many); much more than 1 cm long, or more than 10 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Bark stripping; slash with copious white latex; juvenile leaves lobed; tertiary veins box-like; stipules large; male and female flowers on separate heads on same plant. Taxon images: genarto.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.