Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; compound; pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units present, or absent; specialized inflorescence units umbels; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 0, or 4-6 (sometimes appearing as an untoothed rim); partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-6; free; not fleshy; deciduous; not stalked; entire. Stamens 4-6; filaments present; free of each other; free of the perianth; all equal in size; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary inferior. Carpels joined. Locules 1. Styles absent. Stigmas 1 lobed; distinctive; head shaped. Fruit berry-like; fleshy to non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm; wingless; with folds like marble, in cross- section; aril absent; endospermic.
Compound leaves without stipules; inflorescences terminating in umbels; endosperm ruminate; ovary 1-loculate. Taxon images: arthro.gif arthro2.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.