Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown, black; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked, or four-ranked; not scale-like; simple; of different sorts on a single twig, or all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate, or trinerved, or palmate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible, or not visible; net like, or box like. Intramarginal vein present, or absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 3-5; free; persistent in fruit, or not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3-5; free; not fleshy; deciduous; not stalked; deeply cut in two, or lobed, or fringed like a comb, or entire. Stamens 6, or 8, or 10; filaments present; free of each other; joined to the perianth; all equal in size, or of different sizes; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 3-5. Locules 3-5. Styles free; 3(-5). Stigmas 1 lobed; not distinctive. Fruit berry-like, or drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Leaf axils with 4-6 superimposed serial buds; leaves with 3-5 veins arising from the base; stipules absent; stamens generally unequal and twice as many as petals. Taxon images: genanis.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.